Division of Biology and Medicine
Pathobiology Graduate Program

Courses

Brown University requires a minimum of 24 tuition credits for the PhD degree. Students may continue to register for elective courses that are related to their training throughout their time at Brown.

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Courses

Brown University requires a minimum of 24 tuition credits for the PhD degree. Students may continue to register for elective courses that are related to their training throughout their time at Brown.

Below are suggested courses.  In addition, an entire course catalog is available.

Fall Semester 2025

CodeTitleScheduleInstructor
BIOL 0470GeneticsTuesdays & Thursdays, 10:30 - 11:50amMark Johnson, Erica Larschan & Jody Hall
BIOL 1110Cell Physiology and BiophysicsMondays, 3:00-5:30pmAnita Zimmerman
BIOL 2020Biotechnology Science & IndustryThursdays, 5:00-7:30pmEric Huang
BIOL 1520Innate ImmunityMondays & Wednesdays, 8:30-9:50amLaurent Brossay
BIOL 1535Survey of Biomedical InformaticsMondays, 3:00-5:30pmNeil Sarkar
BIOL 1970AStem Cell BiologyMondays, 2:00-4:30pmRichard Freiman
BIOL 2020Biotechnology Science and IndustryThursdays, 5:00-7:30pmEric Huang
BIOL 2024Gene and RNA Therapeutic DevelopmentTuesdays, 4:30-7:00pmKailene Simon
BIOL 2025Foundations in Statistics for Biology and MedicineTuesdays & Thursdays, 10:30-11:50amLiz Chen
BIOL 2030Foundations for Advanced Study in the Life SciencesTuesdays & Thursdays, 9:00-10:20amGerwald Jogl & Sofia Lizarraga
BIOL 2050Biology of the Eukaryotic CellTuesdays & Thursdays, 1:00-2:20pmChuck Toth
BIOL 2089The Importance of Intellectual Property in BiotechnologyWednesdays, 4:00-6:30pmDaniel Holmander & Jeffrey Morgan
BIOL 2220Topics in Signal TransductionWednesdays, 3:00-5:30pmElena Oancea
BIOL 2222BCurrent Topics in Functional GenomicsTuesdays, 1:00-3:30pmErica Larschan & Carlos Silva-Garcia
BIOL 2250Survey of Modern TherapeuticsTuesdays & Thursdays, 9:00-10:20amDonald Christian
BIOL 2270Advanced BiochemistryTuesdays & Thursdays, 2:30-3:50pmArt Salomon & Juan Alfonzo
BIOL 2300Biomolecular Interactions:  Health, Disease, and Drug designMondays, 3:00- 5:30pmNicolas Fawzi
BIOL 2560Advanced VirologyMondays, Wednesdays, & Fridays, 9:00-9:50am Amanda Jamieson & Sanghyun Lee
BIOL 2860Molecular Mechanisms of DiseaseMondays & Wednesdays, 1:00-2:30pmThomas Bartnikas
ENGN 1945ImmunoengineeringMonday, Wednesday, Friday, 1:00-1:50pmTheresa Raimondo
ENGN 2910SCancer NanotechnologyMonday, Wednesday, Friday, 1:00-1:50pmIan Wong
PHP 2120Introduction to Methods in Epidemiologic ResearchTuesdays & Thursdays, 10:30-11:50amMark Lurie
PHP 2510Principles of Biostatistics and Data AnalysisTuesdays & Thursdays, 9:00-10:20amPeter Lipman
PHP 2560Statistical Programming with RTuesdays & Thursdays, 2:30-3:50pmYing Ma
IMMUN 307QCCancer ImmunologyHarvard Cross-RegistrationKai Wucherpfennig

Spring Semester 2026

CodeTitleScheduleInstructor
BIOL 1250Host-microbiome Interactions in Health and DiseaseThursdays, 2:00 - 4:30pmPeter Belenky & Shipra Vaishnava
BIOL 1295Fundamentals of Cancer ImmunotherapyMondays & Wednesdays, 3:00-4:20pmSean Lawler
BIOL 1550Parasitism:  Biology and DiseaseMondays, Wednesdays, & Fridays, 11:00-11:50amChristopher de Graffenried
BIOL 1555Methods in Informatics and Data Science for HealthMondays, 3:00-5:30pmElizabeth Chen & Neil Sarkar
BIOL 1580Metabolism Across Biological Scales:  Integrative Physiology and PathobiologyTuesdays & Thursdays, 2:30-3:50pmKarthi Chellappa
BIOL 1600Development of Vaccines to Infectious DiseasesMondays & Wednesdays, 3:00-4:20pmShipra Vaishnava & Lalit Beura
BIOL 1820Environmental Health and DiseaseTBAJessica Plavicki
BIOL 2025Foundations in Statistics for Biology and MedicineMondays & Wednesdays, 8:30-9:50amSusannah Colt
BIOL 2040BioimagingMondays, 2:00-5:00pmGeoffrey Williams
BIOL 2167In Vitro Models for DiseaseMondays, 10:00am-12:30pmJacquelyn Schell
BIOL 2170Molecular Pharmacology and PhysiologyMonday, Wednesday, & Fridays, 10:00-11:30amDiana Horrigan
BIOL 2350The Biology of AgingThursdays, 2:00-5:00pmMarc Tatar & Nicola Neretti
BIOL 2525Pathogenomics: Analysis, Interpretation and Applications of Microbial GenomesTuesdays & Thursdays, 10:30-11:50amKatherine Siddle & Christina Cuomo
BIOL 2540Molecular GeneticsTuesdays & Thursdays, 2:30-3:50pmYu-Wen Alvin Huang
BIOL 2865ToxicologyTuesdays & Thursdays, 1:00-2:20pmDaniel Spade
CSCI 2820Algorithmic Foundations of Computational BiologyTuesdays & Thursdays, 2:30-3:50pmSorin Istrail
CLPS 0950Introduction to ProgrammingMondays, Wednesdays, & Fridays, 10:00-10:50amTBD
CLPS 1480BCognitive Aging and DementiaTuesdays & Thursdays, 1:00-2:20pmElena Festa
NEUR 1020Principles of NeurobiologyTuesdays & Thursdays, 9:00-10:20amCarlos Aizenman
NEUR 1540Neurobiology of Learning and MemoryMondays, Wednesdays & Fridays, 10:30-11:50amMonica Linden